Maximum recorded length:
173 mm SL (Murdy, 1989)
230 mm TL (Rainboth, 1996)
Live colouration (Murdy, 1989):
ground colour greenish yellow dorsally and laterally, ventrally greyish, with darkened throat and isthmus; 5 large, midlateral brown spots are usually
visible; often also 5 dark brown saddle-like blotches on dorsum are visible; D1 and D2 membrane yellowish basally, transparent for the rest, with elements
proximally green; caudal fin membrane dusky, with yellow dorsal and ventral margin; anal fin hyaline; pectoral fin base with small red spots, fins proximally
yellowish, with red ventroposterior margin; pelvic fins yellow
Colouration on preservation (Murdy, 1989,
pers. obs.):
ground colour pale brown; blotches and bars frequently retained; D1 and D2 from transparent to dusky; caudal fin dark grey with whitish margins; anal
fin dark grey; pectoral fins medially dusky; pelvic fins hyaline
Diagnosis (Murdy, 1989):
longitudinal scale count 62-81; TRF 20-26; TRB 15-26; predorsal scales 23-38; caudal fin length 19.4-26.7%SL; body depth 10.5-16.5%SL; head depth 9.8-14.4%SL;
head width 9.9-14.4%SL; no black pigment inside mouth; a single large pore posteriorly in interorbital region.
The genus is characterised by a greatly expanded distal head of the fourth epibranchial (see drawing)
Diet:
to the best of my knowledge, the only study available is based on the stomach content of a single Malaysian specimen
(Khaironizam & Norma-Rashid, 2000). This study suggests that its diet
is mainly herbivorous (benthic phytoplankton), including only small amounts (<3%) of animal food items
Reproduction: no published study is available
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